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Resequencing the entire CETP gene uncovers rare variants that independently affect HDL cholesterol in both whites and African blacks (Metabolism 2016)

Original title: Resequencing of the CETP gene in American whites and African blacks: Association of rare and common variants with HDL-cholesterol levels

Metabolism · · 6

Pirim D, Wang X, Niemsiri V, Radwan ZH, Bunker CH, Hokanson JE, Hamman RF, Barmada MM, Demirci FY, Kamboh MI

This study resequenced the entire CETP gene, whose common variants are well documented for their effects on lipid levels and CETP mass/activity, to identify uncommon and rare variants and test their effects on plasma lipid traits, since these effects remained undefined. The gene and flanking regions were resequenced in 190 individuals (95 non-Hispanic whites, 95 African blacks) with extreme HDL-cholesterol levels, identifying 279 sequence variants, 25 of them novel. Selected variants were genotyped in 623 non-Hispanic whites and 788 African blacks, with 184 quality-controlled variants tested via gene-based, single-site, haplotype, and rare-variant (SKAT-O) association analyses. Two novel independent associations, rs1968905 and rs289740, were identified with HDL-cholesterol in African blacks, and rare variants (minor allele frequency below 0.01) were associated with HDL-cholesterol in both non-Hispanic whites (P=0.024) and African blacks (P=0.009).

Read the paper (DOI)PubMed

Original abstract

Background: Cholesteryl ester transfer protein (CETP) plays a crucial role in lipid metabolism. Associations of common CETP variants with variation in plasma lipid levels, and/or CETP mass/activity have been extensively studied and well-documented; however, the effects of uncommon/rare CETP variants on plasma lipid profile remain undefined. Hence, resequencing of the gene in extreme phenotypes and follow-up rare-variant association analyses are essential to fill this gap.

Objective: To identify common and uncommon/rare variants in the CETP gene by resequencing the entire gene and test the effects of both common and uncommon/rare CETP variants on plasma lipid traits in two genetically distinct populations.

Methods And Results: The entire CETP gene plus flanking regions were resequenced in 190 individuals comprising 95 non-Hispanic whites (NHWs) and 95 African blacks with extreme HDL-C levels. A total of 279 sequence variants were identified, of which 25 were novel. Selected variants were genotyped in the entire samples of 623 NHWs and 788 African blacks and 184 QC-passed variants were tested in relation to plasma lipid traits by using gene-based, single-site, haplotype and rare variant association analyses (SKAT-O). Two novel and independent associations of rs1968905 and rs289740 with HDL-C were identified in African blacks. Using SKAT-O analysis, we also identified rare variants with minor allele frequency <0.01 to be associated with HDL-C in both NHWs (P=0.024) and African blacks (P=0.009).

Conclusions: Our results point out that in addition to the common CETP variants, rare genetic variants in the CETP gene also contribute to the phenotypic variation of HDL-C in the general population.

geneticsHDL biology

Summary written by cetpinhibition.org from the published abstract; figures as published. Page updated 19 August 2026. Methods.